Pteropus tokudae (Guam Flying Fox)

[surroundx]

Wiles, Gary J. (1987b). The status of fruit bats on Guam. Pacific Science 41(1-4): 148-157.

[Bhagatí]

Taxonomy, distribution, and natural history of flying foxes (Chiroptera, Pteropodidae) in the Mortlock Islands and Chuuk State, Caroline Islands - Donald W. Buden, Kristofer M. Helgen, Gary J. Wiles
received 20 June 2013 | accepted 17 October 2013 | Published 29 October 2013


Figure 4. Holotype skin of Pteropus tokudae. AMNH 87117, adult male, from Guam, collected 10 Aug 1931 by W.F. Coultas. Scale bar = 25 mm.

source:
www.pensoft.net/J_FILES/1/articles/5840/5840-G-1-layout.html

There is a lot informations about this species.
And bat species of genus Pteropus dwell Mortlock Islands and Chuuk State, Caroline Islands.

ZooKeys 345: 97–135, doi: 10.3897/zookeys.345.5840

[koeiyabe]

"Lost Animals (in Japanese)" by WWF Japan (1996)

[surroundx]

Mickleburgh, Simon P., Hutson, Anthony M. and Racey, Paul A. (1992). Old World fruit bats. An action plan for their conservation. Gland, Switzerland: IUCN. viii + 252 pp.

[Sebbe]

Each flying fox on its own branch: A phylogenetic tree for Pteropus and related genera (Chiroptera: Pteropodidae)

Pteropodidae is a diverse Old World family of non-echolocating, frugivorous and nectarivorous bats that includes the flying foxes (genus Pteropus) and allied genera. The subfamily Pteropodinae includes the largest living bats and is distributed across an immense geographic range from islands in East Africa to the Cook Islands of Polynesia. These bats are keystone species in their ecosystems and some carry zoonotic diseases that are increasingly a focus of interest in biomedical research. Here we present a comprehensive phylogeny for pteropodines focused on Pteropus. The analyses included 50 of the ∼63 species of Pteropus and 11 species from 7 related genera. We obtained sequences of the cytochrome b and the 12S rRNA mitochondrial genes for all species and sequences of the nuclear RAG1, vWF, and BRCA1 genes for a subsample of taxa. Some of the sequences of Pteropus were obtained from skin biopsies of museum specimens including that of an extinct species, P. tokudae. The resulting trees recovered Pteropus as monophyletic, although further work is needed to determine whether P. personatus belongs in the genus. Monophyly of the majority of traditionally-recognized Pteropus species groups was rejected, but statistical support was strong for several clades on which we based a new classification of the Pteropus species into 13 species groups. Other noteworthy results emerged regarding species status of several problematic taxa, including recognition of P. capistratus and P. ennisae as distinct species, paraphyly of the P. hypomelanus complex, and conspecific status of P. pelewensis pelewensis and P. p. yapensis. Relationships among the pteropodine genera were not completely resolved with the current dataset. Divergence time analysis suggests that Pteropus originated in the Miocene and that two independent bursts of diversification occurred in the Pleistocene in different regions of the Indo-Pacific realm.

www.sciencedirect.com/science/article/pii/S1055790314001092

[Sebbe]

Removal of 21 Species From the List of Endangered and Threatened Wildlife.

By 1945, fruit bats were difficult to locate even in the northern half of Guam, where they were largely confined to forested cliff lines along the coasts (Baker 1948, p. 54). During surveys conducted between 1963 and 1968, the Guam Division of Aquatic and Wildlife Resources (DAWR) confirmed that bats were declining across much of Guam and were absent in the south. It was also during these same field studies that the third and last little Mariana fruit bat was collected in northern Guam in 1968 (Baker 1948, p. 146). Increased survey efforts during the late 1970s and early 1980s reported no confirmed sightings of the little Mariana fruit bat (Wheeler and Aguon 1978, entire; Wheeler 1979, entire; Wiles 1987, entire; Wiles 1987, pp. 153–154). In the final rule listing the little Mariana fruit bat as endangered (49 FR 33881; August 27, 1984), we noted that the species was on the verge of extinction and had not been verifiably observed after 1968. When we published a joint recovery plan for the little Mariana fruit bat and the Mariana fruit bat in 1990, we considered the little Mariana fruit bat already extinct based upon the available literature (USFWS 1990, p. 7). During the 1990s, Mariana fruit bat numbers on Guam decreased and fatalities of immature bats increased, hypothesized to be a result of predation by the brown tree snake (Wiles et al. 1995, pp. 33–34, 39–42). With bat abundance continuing to decline in the 2000s, the island’s Mariana fruit bat population currently fluctuates between 15 and 45 individuals (Mildenstein and Johnson 2017, p. 24; USFWS 2017, p. 54). Even if the little Mariana fruit bat persisted at undetectable numbers for some time after its last confirmed collection in 1968, it is highly likely the little Mariana fruit bat experienced the same pattern of decline that we are now seeing in the Mariana fruit bat. Time Since Last Detection As stated above, the little Mariana fruit bat was last collected in northern Guam in 1968 (Baker 1948, p. 146). Intensive survey efforts conducted by Guam DAWR and other researchers in subsequent decades have failed to locate the species. Decades of monthly (and, later, annual) surveys for the related Mariana fruit bat by qualified personnel in northern Guam have failed to detect the little Mariana fruit bat (Wheeler and Aguon 1978, entire; Wheeler 1979, entire; Wiles 1987, entire; Wiles 1987, pp. 153–154; USFWS 1990, p. 7).

www.govinfo.gov/content/pkg/FR-2023-10-17/pdf/2023-22377.pdf